Introduction 1
Material and Methods 6
Characteristics of Epiphyllous Liverworts and Adaptations to the
Phyllosphere 9
Growtth habit 9
Stem structure 9
Leaves 10
Leaf cells 10
Oil bodies 12
Ocellus 14
Perianth 16
Spores 16
Asexual reproductive organs 19
Neotenic plants 21
Rhizoids 21
Types of Epiphyllous Liverworts 22
Relationship between Epiphyllous Liverworts and Host Leaves 23
Diversity of Epiphyllous Liverworts 24
Ecology and Distribution of Epiphyllous Liverworts 27
Conservation of Epiphyllous Liverworts 29
Taxonomic Treatment 30
Systematic arrangement of Chinese epiphyllous liverworts 30
Key to epiphyllous families 32
I. METZGERIACEAE 32
Metzgeria Raddi 32
II. JUNGERMANNIACEAE 37
Jamesoniella (Spruce) F.Lees 37
III. PLAGIOCHILACEAE 38
Plagiochila (rumort.) Dumort 39
IV. GEOCALYCACEAE 45
Heteroscyphus Schiffn 45
V. CEPHALOZIACEAE 51
Odontoschisma (rumort.) Dumort 52
VI. CALYPOGEIACEAE 53
Calypogeia Raddi 53
VII. RADULACEAE 55
Radula Dumort 56
VIII. FRULLANIACEAE 72
Frullania Raddi 72
IX. JUBIULACEAE 82
Jubula Dumort 82
X. LEJEUNEACEAE 85
Key to genera and subfamilies of epiphyllous Lejeuneaceae 85
Trocholejeunea Schiffn 86
Ptychanthus Nees 88
Spruceanthus Verd 91
Lopholejeunea (Spruce) Schiffn 92
Caudalejeunea (Steph.) Schiffn 96
Leucolejeunea A. Evans 101
Cheilolejounea (Spruce) Schiffn 104
Lepidolejeunea R.M.Schust 124
Lejeunea Lib 126
Stenolejeunea R.M.Schust 162
Metalejeunea Grolle 166
Otolejeunea Grolle & Tixier 168
Drepanolejeunea (Spruce) Schiffn 169
Leptolejeunea (Spruce) Schiffn 200
Tuyamaella S.Hatt 218
Diplasiolejeunea (Spruce) Schiffn 224
Colura (rumort.) Dumort 232
Cololejeunea (Spruce) Schiffn 247
Subg. Pedinolejeunea Benedix ex Mizut 248
Subg. Cryptolejeunea Benedix 282
Subg. Leptocolea (Spruce) Schiffn 283
Subg. Cololejeunea 311
Subg. Tacniolejeunea (Zwickt) Benedix 340
Aphanolejeunea A.Evans 368
Dubia and Excludenda 375
Taxa Not Available for the Present Study 379
Acknowledgements 383
Literature Cited 384
Glossary 404
Index to Scientific Names 407